The Recently Extinct Plants and Animals Database Extinct Mammals: Marsupials: Thylacinus cynocephalus

Scientific Name: Thylacinus cynocephalus

Species Authority: Harris, 1808:174

Synonym/s: Didelphis cynocephala Harris, 1808:174 (original combination); Thylacinus harrisii Temminck, 1824; Dasyurus leucocephalus Grant, 1831; Thylacinus striatus Warlow, 1833; Thylacinus communis Anon, 1859; Thylacinus breviceps Krefft, 1868:296; Thylacinus spelaeus Owen, 1845; Thylacinus major Owen, 1877; Thylacinus rostralis De Vis, 1894

Common Name: Thylacinus, Tiger-wolf, bull dog thylacine (Krefft re. T. breviceps); for a complete list refer to (Guiler & Godard, 1998:15)

Locality: Australia (including Tasmania) and New Guinea

Last Record: 1933 (wild) (Pearse, 1976); 7 September 1936 (captivity) (Guiler, 1985)

IUCN status: Extinct

TSE forum thread:

The Museum of Comparative Zoology, Harvard, has three specimens:

MCZ 6014 (mounted skeleton; sex unspecified)
MCZ 6349 (mounted skin; sex unspecified)
MCZ 36797 (most elements present; sex unspecified)

Merseyside County Museums, Liverpool, have two specimens:

MCM 26.9.1910 (ss and limb bones) (Fisher, 1984:208)
MCM 1979.21 (sk) (Fisher, 1984:208)

The Hunterian Museum and Art Gallery, University of Glasgow, has two specimens:

GLAHM Z503 (mount, sex not specified, see here)
GLAHM Z1358 (skull, juvenile female, see here)

The Grant Museum of Zoology has 6 specimens:

Z87 (skull, lacking lower mandible; female)
Z88 (skull and mandible)
Z89 (mounted skeleton)
Z90 (skull and lower mandible)
Z1479 (skull and mandible; male)
Z1653 (four fluid body part specimens)

South Australian Museum

SAM M665/001
SAM M922
SAM M1952-56
SAM M1959-60

Zoology Museum, Ghent University (Belgium)

Has one specimen, probably from the collection of C. J. Temminck (Verschelde & Adriaens, n.d.).

Other specimens:

S 1646
NMSZ 1980.67 (adult female)
NMSZ 1868.30.1 (adult male)
WAM F6353
WAM F6358
WA M33 (Kitchener & Vicker, 1981:156)
WA M195 (Kitchener & Vicker, 1981:156)
WA M3318 (Kitchener & Vicker, 1981:156)
WA M17189 (Kitchener & Vicker, 1981:156)
USNM 125345 (skin, adult male)
USNM 49723
USNM 49724 (skull of USNM 125345, adult male) (Senter & Moch, 2015)
USNM 155387 (Senter & Moch, 2015)
USNM 238801 (Senter & Moch, 2015)
NRM 566599 (ethanol preserved, in Sweden)
NRM 592206 (mounted skin, adult male)
NMV C5752 (alcohol-preserved skull)
NMV C5748
NMV C5747
NMV P187757
QMF44643 ("I1")
QM F726 in part (Mackness et al. 2002:238; Louys & Price, 201X:21)
QM F3744 (Louys & Price, 201X:21)
AMNH 160248 (van Deusen, 1963:280)
AMNH 77701 (van Deusen, 1963:280)
AMNH 35866 (Mittelbach & Crewdson, 2006)
NMA 1984.0010.0706 (preserved pouch and scrotal sac) (Sleightholme, 2011:954)
The Natural History Museum, Dublin has four thylacines (source), including NH:1917.25.1 (source)
NH:1917.25.1 (national Museum of Ireland, source)

The International Thylacine Specimen Database (ITSD)

The ITSD is a database of all known preserved thylacine specimens (subfossils are not included at present) held in museum collections around the world; and in 9 further instances, in private collections as well. The latest, revised edition (i.e. 5th) of the database is available as a CD (as were the others); however, they are only available to scientists and thus the public is deprived of much valuable information. But a basic breakdown of the database can be found online. A sixth revision is planned for 2017 (Sleightholme & Campbell, In Press).

Attacks on Humans

"A curious circumstance happened at Mr. Blinkworth's, Jerusalem, the other day. A native tiger, as it is called, boldly entered his cottage, where his family was assembled, and seized one of the little children by the hair, but fortunately missed its bite. Mr. Blinkworth who was confined to the house with a lame hand, alertly seized the animal by the tail and dashing it on the ground speedily killed it."

(Source: Hobart Town Courier, Saturday, 17 April, 1830)

The Scottsdale North-Eastern Advertiser reported an attack on a Miss Priscilla Murray by a one-eyed Thylacine in 1900 (Smith, 1982:249).

The Sydney Morning Herald (22 May, 1872) reported that a Mr James Jones was approached by a tiger coming out of the scrub (quoted by Whitley, 1973).

Data Sets of Sightings

Smith (1981) and Rounsevell & Smith (1982) both dealt with sightings in Tasmania to 1978. The latter analysed 104 sightings during the period 1970-1979, and stated that 84 sightings had been reported during the period 1960-1969.

Heberle (2004) analysed 203 reported sightings from Western Australia between 1936 and 1998.

According to the Australian Rare Fauna Research Association (ARFRA), they have "some 3800 mainland sightings of an animal answering the description of the Tassie Tiger", as well as documentation of "predation, vocalisations and prints".

Murray McAllister provides anecdotes and reports sent to him by many different people who claim to have seen the thylacine:

Well-known Australian zoologist Gary Opit has recorded 64 possible thylacine sightings as a result of people calling in during his radio show:

A whole batch of reports have been officially released by the Tasmanian government:

This is interesting:

"According to the original commentary for the DVD of the Howling 3 (2002-2007 edition, later versions lack the commentary) the director originally wanted a thylacine scene so made up a dog to resemble that animal. Before shooting, however, the animal escaped and several mainland thylacine sightings were the result.

It may be informative to find out the date and location of that movie[']s shooting to see if any reports date from that location and time. If so those reports can probably be eliminated."


"My Dad once told me (and my mother) that he had seen a live one at Healesville Sanctuary (Vic) when he was a boy, in the late 1920's or early 30's.

We were talking about this recently and she rang the Sanctuary and checked, and yes they admitted they did have a couple of Tigers in captivity there but they don't like to admit they actually had them!!!"



The 1996 documentary, still the best to date:

A Wilf Batty interview

An interview with Alison Reid, who should need no introduction ;)

An Animal X series segment on cloning the thylacine:

An Animal X series episode containing a story about mainland thylacine sightings/encoutners:

Cloning the Tasmanian Tiger, a documentary about the work of Dr. Mike Archer and his team in trying to bring the Thylacine back to life.

The Monster Quest episode on the Tasmanian tiger, including interviews with Col Bailey and other "believers".

Murray McCalister, Victorian thylacines and headless 'roos, including eyewitness reports

An episode of The Big Country television series about Jeremy Griffith's search for the thylacine (1973):

The Animal X TV episode on the thylacine, narrated by the late Bill Kerr:

The only 19th century photograph of a living thylacine:

"Timid Devils and Ghost Tigers"

Robert Bagshaw's search for the thylacine

Reconstruction of the thylacine's running gait, contrast with the "Doyle footage":

Fictional web series (Out of the Shadows) concerning the Tasmanian tiger:

A children's program that fuses fact and fiction:

The brief segment on the thylacine, part of David Day's television program:

Below is an incomplete bibliography on the thylacine. The section on scientific papers is relatively complete except for the 19th century period, however the section on popular articles, especially newspaper stories, is far from complete. I have started compiling a list of thylacine articles available through Trove (see here), which will take some time to complete. In addition, many local newspapers occasionally carry stories of reports of the thylacine, which probably number in the thousands by now.

Original Scientific Description:

Harris, George Prideaux. (1808). Description of two new species of Didelphis from Van Diemen's Land. Transactions of the Linnean Society of London 9(1): 174-178.

Other references:

Berry, Colin. (1997). The Tasmanian Tiger Trail. Wynyard, Tasmania: Self published. 14 pp.

Brodie, Jedediah F. et al. (2017). Crossing the (Wallace) line: local abundance and distribution of mammals across biogeographic barriers. Biotropica. DOI: 10.1111/btp.12485 [Abstract]

Camens, Aaron B., Carey, Stephen P. and Arnold, Lee J. (2017). Vertebrate trace fossils from the Late Pleistocene of Kangaroo Island, South Australia. Ichnos: A Journal for Plant and Animal Traces. [Abstract]

Jones, Neil. (2003). The Kenilworth dasyuroid: the Tasmanian Tiger has relatives spotted in Queensland. Kenilworth, Queensland: Self published. 16 pp.

Lawton, Graham. (2017). Eye on the tiger. New Scientist 234(3124): 40-41. [Abstract]

Lindeen, Carol. (2007). Tasmanian tiger: Thylacine cynocephalus. Mankato, Minn.: Capstone Press.

Marks, Clive A. et al. (2017). Trends in anecdotal fox sightings in Tasmania accounted for by psychological factors. Conservation Biology. DOI: 10.1111/cobi.12944

Morrell, Sascha. (2017). Thylacine. Meanjin 76(1): 189. [Abstract]

Pyers, Greg. (2007). Finding Out About Thylacines. Port Melbourne, Victoria: Echidna Books. 32 pp.

Smith, A. (2017). Estimating odds against spotting a thylacine. New Scientist 234(3127): 54.

Ackman, Phil. (1976). Wild dog pack could be 'extinct' native tigers. Sunday Telegraph, 8 August, p. 5.

Australian Geographic. (Due October, 2017). Australia Gone Wild. Sydney: Australian Geographic. [Available from Andrew Isles NHB]

Haygarth, Nic. (2017). The myth of the dedicated thylacine hunter: Stockman-hunter culture and the decline of the thylacine (Tasmanian tiger) in Tasmania during the late Nineteenth and early Twentieth centuries. Papers and Proceedings: Tasmanian Historical Research Association 64(2): 30-45. [Abstract]

The Wonder Book of Empire, p. 135. (

Hoare, Philip. (2014). The Sea Inside. London: Harper Collins Publishers.

Smith, Vivian. (1956). The Other Meaning. Publisher? [a book of poetry, which includes a poem on the thylacine]

Williams, Michael "Mike". (2014). The Truth About the Nullarbor Thylacine, pp. 129-140. In: Lang, Rebecca (ed.). The Tasmanian Tiger: Extinct or Extant? Hazelbrook, NSW: Strange Nation Publishing. 186 pp.

Clark, Jerome. (2013). Unexplained!: Strange Sightings, Incredible Occurrences, and Puzzling Physical Phenomena, third edition. Canton, Michigan: Visible Ink Press. [chapter title: Thylacines, pp. 198-208]

Strahan, Ronald. (1986). [letter to the editor], 26 June, p. 82.

The Sydney Mornin Herald, 29 July, 1986, "Colour Photos of Tasmanian Tiger Doubted" [contains an interview with Athol Douglas; fide (Williams, 2014:136)]

Carlson, Colin J., Bond, Alexander, L. and Burgio, Kevin R. (Accepted, 2017). Estimating the extinction date of the thylacine with mixed certainty data. Conservation Biology. DOI: 10.1111/cobi.13037 [Abstract]

Archer, Michael "Mike". (1978). Quaternary vertebrate faunas from the Texas Caves of southeastern Queensland. Memoirs of the Queensland Museum 19(1): 61-109.

M. J. Tyler. 1977. Pleistocene frogs from caves at Naracoorte, South Australia. Transactions of the Royal Society of South Australia 101(3):85-89.

G. J. Prideaux. 2004. Systematics and evolution of the sthenurine kangaroos. In S. W. Awramik, A. Barnosky, J. A. Doyle, M. L. Droser, P. M. Sadler (eds.), UC Publications in Geological Sciences, University of California Press 146:1-623.

J. H. Hope and H. E. Wilkinson. 1982. Warendja wakefieldi, a new genus of wombat (Maruspialia , Vombatidae) from Pleistocene sediments in McEacherns Cave, western Victoria. Memoirs of the National Museum of Victoria 43:109-120.

D. Merrilees. 1973. Fossiliferous deposits at Lake Tandou, New South Wales, Australia. Memoirs of the National Museum of Victoria 34:177-182.

R. E. Molnar, L. S. Hall, and J. H. Mahoney. 1984. New fossil localities for Macroderma Miller, 1906 (Chiroptera, Megadermatidae) in New South Wales and its past and present distribution in Australia. Australian Mammalogy 7(1-2): 63-73.

R. A. Fraser and R. T. Wells. 2006. Palaeontological excavation and taphonomic investigation of the late Pleistocene fossil deposit in Grant Hall, Victoria Fossil Cave, Naracoorte, South Australia. Alcheringa: An Australasian Journal of Palaeontology 30(S1): 147-161.

Le Guellec-Minel, Anne. (2018). The Tasmanian Tiger: From Extinction to Identity Myth in White Australian Society and Fiction, pp. 67-83. In: Dodeman, André and Raimbault, Élodie (eds.). Literary Location and Dislocation of Myth in the Post/Colonial Anglophone World. Leiden, The Netherlands: Koninklijke Brill NV.

Mooney, Nick. (2016). Thylacine; the Improbable Tiger. Capeia: 20160907.002.

Sleightholme, Stephen R. and Campbell, Cameron R. (In Press, 2017). The International Thylacine Specimen Database (6th Revision - Project Summary & Final Report). Australian Zoologist. [Abstract]

Dando, W. P. (c.1920). More Wild Animals & The Camera. Jarrold & Sons. [contains a photograph of the thylacine]

Woolley, P. A. (1987). The seminiferous tubules, rete testis and efferent ducts in didelphid, caenolestid and microbiotheriid marsupials, pp. 217-227. In: Archer, Michael (ed.). Possums and Opossums: Studies in Evolution. Sydney, N.S.W.: Surrey Beatty & Sons / Royal Zoological Society of New South Wales.

Puxley, W. L. (1949). Strayed Among Lonely Islands. London: Williams & Norgate.

Block, Erik. (2017). The History of the Thylacine in Captivity. Zoo Grapevine & International Zoo News 44: 29-33.

Chapple, Peter. (2001). The Quest for the Thylacine, pp. 75-81. In: Cropper, Paul (ed.). Myths & Monsters 2001 Conference Papers. Unpublished.

Anonymous. (2001). The Mainland Thylacine: An overview, pp. 82-83. In: Cropper, Paul (ed.). Myths & Monsters 2001 Conference Papers. Unpublished.

White, Lauren C., Mitchell, Kieren J. and Austin, Jeremy J. (2017). Ancient mitochondrial genomes reveal the demographic history and phylogeography of the extinct, enigmatic thylacine (Thylacinus cynocephalus). Journal of Biogeography. DOI: 10.1111/jbi.13101 [Abstract]

Kealy, S. and Beck, R. M. D. (In Press, 2017). Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology. [Abstract]

Turnbull, Paul. (2017). Science, Museums and Collecting the Indigenous Dead in Colonial Australia (Palgrave Studies in Pacific History book series). Cham: Palgrave Macmillan. [chapter 2: Skeletal Collecting Before Darwin, pp. 71-96]

Thorn, Kailah M. et al. (2017). Fossil mammals of Caladenia Cave, northern Swan Coastal Plain, south-western Australia. Records of the Western Australian Museum 32: 217-236.

Feigin, Charles Y. et al. (2017). Genome of the Tasmanian tiger provides insights into the evolution and demography of an extinct marsupial carnivore. Nature Ecology & Evolution. doi:10.1038/s41559-017-0417-y

Data from: Letting the “cat” out of the bag: pouch young development of the extinct Tasmanian tiger revealed by X-ray computed tomography

White, Lauren C., Saltré, Frédérik, Bradshaw, Corey J. A. and Austin, Jeremy J. (2018). High-quality fossil dates support a synchronous, Late Holocene extinction of devils and thylacines in mainland Australia. Biology Letters 14: 20170642.

Dunnet, G. M. and Mardon, D. K. (1974). A monograph of Australian fleas (Siphonaptera). Australian Journal of Zoology, Supplementary Series No. 30: 1-273, 366 illus., maps. [volume 22?] [Uropsylla tasmanica recorded from the thylacine]

Barnett, Ross and Lorenzen, Eline. (2018). Thylacine tales. Nature Ecology & Evolution 2: 7-8.

Smith, G. Elliot. (1902). On a peculiarity of the cerebral commissures in certain Marsupialia, not hitherto recognised as a distinctive feature or the Diprotodontia. Proc. R. Soc. Lond. 70: 226-231.

Parnaby, Harry, Ingleby, Sandy and Divljan, Anja. (2017). Type specimens of non-fossil mammals in the Australian Museum, Sydney. Records of the Australian Museum 69(5): 277-420.

Jackson, S. M., and C. P. Groves. 2015. Taxonomy of Australian Mammals. Melbourne, Victoria: CSIRO Publishing. 529 pp.

Newton, Axel H. et al. (2018). Letting the ‘cat’ out of the bag: pouch young development of the extinct Tasmanian tiger revealed by X-ray computed tomography. R. Soc. open sci. 5: 171914.

References A-D
Species Bibliography:

ABC News “Tasmanian tiger DNA comes alive in a mouse,”. 5202008.

Adam-Smith, Patsy. (1968). Tiger Country: Remote Parts of Tasmania. Sydney: Rigby Ltd.

Adam-Smith, Patsy. (1975). Tiger Country: Remote Parts of Tasmania. Publisher?

Agnone, Julie Vosburgh. (1998). Any sign of a thylacine? National Geographic World 269: 5.

Allport, M. 1868. Remarks on Mr. Krefft’s “Notes on the Fauna of Tasmania”. Papers and Proceedings of the Royal Society of Tasmania: 33-36.

Anderson, Alistair. (2016). Exploring the Affective (After)Lives of Digital Archives. Bristol Society and Space (blog), 1 June, available at:

Andrews, A. P. (c.1975). Thylacine Thylacinus cynocephalus (Tasmanian tiger, marsupial wolf). Education Leaflet No. 8. Hobart: Tasmanian Museum and Art Gallery.

Andrews, A. P. (1985). Thylacine. Thylacinus cynocephalus. Pamphlet produced by the Tasmanian Museum and Art Gallery, Macquarie Street, Hobart, Tasmania.

Andrews, Crispin. The Tasmanian tiger meets the British lion: [Did a thylacine ever lurk in the English Lake District?] [online]. 40 [degrees] South, No. 63, Summer 2011-2012: 74-76.

Andrews, Crispin and Lang, Rebecca. (2016). Tiger time. 40 [degrees] South 82: 61-63. [Abstract]

Anonymous. (1805). ["An animal of a truly singular and nouvel description..."]. Sydney gazette and NSW Advertiser vol. 3, no. 112, 21 April.

Anonymous. (1823). ["article on a thylacine incident"]. Hobart Town Gazette, 2 August, p. 2.

Anonymous. (1884). The Mercury, 26 August 1884, p. 2.

Anonymous. (1885a). The Tasmanian, 13 June 1885.

Anonymous. (1885b). The Tasmanian, 11 July 1885.

Anonymous. (1886). The Mercury, 8 October 1886.

Anonymous. (1887a). The Tasmanian Mail, Supplement, 30 April 1887.

Anonymous. (1887b). The Tasmanian News, 6 September 1887.

Anonymous. (1891). Hobart Mercury, 5 December 1891.

Anonymous. (1893). Hobart Mercury, 9 May 1893.

Anonymous. (1895). The Tantanoola "Tiger". The Advertiser, Monday 22 July. [It is suggested that the Tantanoola Tiger is a thylacine]

Anonymous. (1897). Tasmanian tigers. Agricultural Gazette, June, p. 178-179.

Anonymous. (1899). Launceston Examiner, 22 March 1899.

Anonymous. (1911). Weekly Courier, 7 December 1911.

Anonymous. (1924). The Mercury, 70th Anniversary Special Issue, supplement: 'Preserving Our Native Animals and Birds: Phenomenal Success of the Beaumaris Zoo', 5 July 1924.

Anonymous. (1937a). Tasmanian Tigers: Evidence of Existence on West Coast. Examiner, Friday 19 February.

Anonymous. (1937b). Addition to Zoo Banned. The West Australian, Wednesday 24 March. [Thylacines afforded protection, making it illegal to capture or kill them]

Anonymous. (1945). Permit to Hunt Tasmanian Tigers. Townsville Daily Bulletin, 13 August.

Anonymous. (1953). Thylacine Tracks Found, Cast. The Mercury, Monday 12 January.

Anonymous. (1963). Animal in Danger. Animals 2(2): pagination?. [J. H. Calaby: "Photograph inside front cover and note on Thylacinus"]

Anonymous. (1966a). Rare tiger trace seen. Science News, 20 August, p. 118. [volume 90, issue 20?]

Anonymous. (1966b). Thylacine in Western Australia. Monthly Service Bulletin 15: 118. [some kind of publication authored by a staff member of CALM, the predecessor of the DEC]

Anonymous. (1969a). Tigers, Devils, monsters and things that walk in the night. Wildl. in Aust. 6: 54.

Anonymous. (1969b). The Thylacine or "Tasmanian Wolf". Leaflet (Australian Museum), no. 49. Sydney: Australian Museum.

Anonymous. (1970). The rare thylacine. Nature Walkabout 6(2): 5-7.

Anonymous. (1971). Reader remembers the "hyenas". Burnie Advocate, 8 May.

Anonymous. (1973a). Another strange beast sighted on Tableland. Cairns Post, 17 Jan.

Anonymous. (1973b). Aloomba woman reports seeing Tableland beast. Cairns Post, 18 Jan.

Anonymous. (1974a). Paintings of Tas. Tiger found in NT. Sydney Morning Herald, 27 April.

Anonymous. (1974b). The Northern Territory's prehistoric monster (tiger?). Sydney Morning Herald, 6 May.

Anonymous. (1977a). 'Tiger' now in baby mystery. Sunday Telegraph (Sydney), 27 March, p. 46.

Anonymous. (1977b). 'Extinct' Tiger seen claim two. Sunday Telegraph (Sydney), 21 August.

Anonymous. (1977). Thylacine. 6: 94–95 in The Australian Encyclopaedia. Vol. 4. Grolier Society of Australia : Sydney 3rd edn.

Anonymous. (1979a). Naturalists hunt extinct tiger in Tasmania. The Times (London), 16 November.

Anonymous. (1979b). "The Mercury" (6/12/77), p. 3, Hobart.

Anonymous. (1980a). The $55,000 search to find a Tasmanian tiger. The Australian Women's Weekly 24 Sept., pp. 42-43.

Anonymous. (1980b). Tasmanian Tiger eludes search. Hong Kong Standard, 21 September.

Anonymous. (1980c). Tasmanian tiger sought in Australia. Washington Post, 29 May.

Anonymous. (1981). Search on for tiger with a pouch. Pretoria News, 24 October, p. 8.

Anonymous. (1982). Eighty-year-old Tasmanian Tiger dissected. Omega Science Digest 1982(July/August): 27.

Anonymous. (1984a). Last gape of the Tasmanian tiger. Nature 307: 411.

Anonymous. (1984b). Last gape of the Tasmanian tiger, p.76-77. In: Beyond Vision. Oxford, Oxford University Press.

Anonymous. (1984c). Tassie tiger's tale. [publication?] 7(4): 4-5.

Anonymous. (1988). Where's that tiger? National Geographic World 152: 34.

Anonymous. (2002a). The Mercury, 2 June 2002.

Anonymous. (2002b). Thylacine reborn? Earth Island Journal 17(3): 18-19.

Anonymous. (2003). Thyla seen near CBD? Sydney Morning Herald, 18 August.

Anonymous. (2013). Tassie tigers in the area? Casey Weekly Cranbourne, 27 January. [a previous story elicited to new sightings to be reported to Michael Moss]

Anonymous. (2014). Novel take on 'extinct' tiger. The Herald (Scotland), Friday 8 August. [author of a thylacine novel to search for the animal in Tasmania]

Archer, Michael. (1971). A re-evaluation of the Fromm's Landing Thylacine tooth. Proceedings of the Royal Society of Victoria 84: 229-234.

Archer, Michael. (1974). New information about the Quaternary distribution of the Thylacine (Marsupialia: Thylacinidae) in Australia. Journal of the Royal Society of Western Australia 57(2): 43-50.

Archer, Michael. (1976). The dasyurid dentition and its relationships to that of didelphids, thylacinids, borhyaenids, (Marsupicarnivora) and peramelids (Peramelina; Marsupialia). Australian Journal of Zoology, Supplementary Series 24(39): 1-34. [Abstract]

Archer, Michael. (1979). The status of Australian dasyurids, thylacinids and myrmecobiids, pp. 29-43. In: Tyler, Michael J. (ed.). The Status of Endangered Australasian Wildlife. Proceedings of the Centenary Symposium of the Royal Zoological Society of South Australia, Aselaide, 21-23 September, 1978. [pagination is taken from the 1980 reprint, may actually be pp. 23-42?]

Archer, M., 1982. Thylacinus cynocephalus (Harris, 1808). Pp. 91-93 in "The red data book". I.U.C.N.: Switzerland.

Archer, M. (1984a). The status of Australian dasyurids, thylacinids and myrmecobiids, pp. 1015-21. In: Archer, M. and Clayton, G. (eds). Vertebrate zoogeography and evolution in Australasia (Animals in space and time). Carlisle, Western Australia: Hesperian Press.

Archer, Michael. (1984b). Doubts on Thylacine. Aust. Nat. Hist. 21: 263.

Archer, Michael. (1987). A wolf in kangaroo's clothing. Pp. 70-72 in "The Antipodean ark" ed S. Hand, M. Archer. Angus & Robertson Publishers: Sydney. [relevant citation?]

Archer, Michael. (1993). The Murgon monster. Australian Natural History 24(4): 60-61.

Archer, Michael. (1997). Tiger, tiger out of sight. Nature Australia 25(8): 70-71.

Archer, Michael. (2003). Cloning the Thylacine: the “yes” case. 40° South Tasmania 28: 19-20.

Archer, Mike. (year?). “Not dead, just stuffed,”. The Bulletin 3232005. 123 (13):

Archer, M. and Baynes, A. (1972). Prehistoric mammal faunas from two small caves in the extreme south-west of Western Australia. Journal of the Royal Society of Western Australia 55: 80-89.

Archer, M., Clayton, G. and Hand, S. J. (1984). A checklist of Australasian fossil mammals, pp. 1027-1087. In: Archer, M and Clayton, G. (eds.). Vertebrate Zoogeography and Evolution in Australasia: (Animals in space and time). Carlisle, Western Australian: Hesperian Press.

Archer, Michael, Hand, S. J. and Godthelp, Hank. (1991). Riversleigh: The Story of Animals in Ancient Rainforests of Inland Australia. Sydney: Reed Books.

Arredondo, Oscar. (1981). Reemplazo de Paracyon por Indocyon (Carnivora: Canidae). Misc. Zool. Acad. Cien. Cuba 12: 4.

Ashwell, K. W. (2008). Encephalization of Australian and New Guinean Marsupials. Brain, Behavior and Evolution 71(3): 181-199. [Abstract]

Attard, Marie R. G. (2012). Unveiling the mysteries of the Tasmanian Tiger. The conversation.

Attard, Marie R. (2013). Who’s on the menu: Diet and extinction risk of the thylacine. Ph.D. thesis. Biological Sciences, School of Biological, Earth and Environmental Sciences, University of New South Wales, Australia.

Attard, Marie R. Chamoli, U. Ferrara, T. L. Rogers, T. L. Wroe, S. (2011). Skull mechanics and implications for feeding behaviour in a large marsupial carnivore guild: the thylacine, Tasmanian devil and spotted-tailed quoll. Journal of Zoology 285(4): 292-300.

Attard, M. R. G., Parr, W. C. H., Wilson, L. A. B., Archer, M., Hand, S. J. et al. (2014). Virtual Reconstruction and Prey Size Preference in the Mid Cenozoic Thylacinid, Nimbacinus dicksoni (Thylacinidae, Marsupialia). PLoS ONE 9(4): e93088.

Attard, Marie R. G. and Wroe, Stephen. (2012). The Thylacine Myth. Australasian Science 33(5): 19-22.

Attard, M. R. G., Wroe, S. and Rogers, T. L. (In prep) Who’s on the menu? Stable isotopes reveal the thylacine’s diet and potential for competition.

Australian Museum. (1964). The Thylacine or 'Tasmanian Wolf'. Leaflet No. 49. Sydney: V. C. N. Blight.

Australian Museum. (1991). Catalogue of Thylacine Specimens. Sydney: Mammal Department, Australian Museum.

Australian Museum. (2000). The Australian Museum Rheuben Griffiths Trust Thylacine Project. Sydney : The Australian Museum. 14 pp. ill.

Australian National Parks and Wildlife Service. (1978). Thylacine, Thylacinus cynocephalus. Rare and Endangered Species Leaflet. Mammals No. 9. Canberra: Australian Government Publishing Service.

Author?. (1880). Guide alle Gallerie del Museo Zoologico e di Anatomia Comparata di Torino. Torino: Stamperia Reale di Torino. Small 8vo, 30pp, ads [44].

Author?. (1997). Here, There Everywhere! The Australian Magazine 1997(November 15-16): 12-18.

Author?. (2016). The Thylacine — or Tasmanian Tiger. Cryptid Culture 3: 50-53.

Ayliffe, L. K., G. J. Prideaux, M. I. Bird, R. Grün, R. G. Roberts, G. A. Gully, R. Jones, L. K. Fifield, and R. G. Cresswell. 2008. Age constraints on Pleistocene megafauna at Tight Entrance Cave in southwestern Australia. Quaternary Science Reviews 27: 1784-1788.

B. A. N. I. (1946). Puzzle of the Tasmanian tiger. Country Life, 23 August, p. 355.

Bagust, Phil. (2006). Vampire Dogs and Marsupial Hyenas: Fear, Myth, and the Tasmanian Tiger’s Extinction, pp. 93-105. In: Day, Peter (ed.). Vampires: Myths and Metaphors of Enduring Evil. Amsterdam and New York: Rodopi.

Bailey, Col. (2001). Tiger Tales: Stories of the Tasmanian Tiger. Sydney: HarperCollins. xii + 164 pp.

Bailey, Col. (2013). Shadow of the Thylacine: One Man's Epic Search for the Tasmanian Tiger. Scoresby: Five Mile Press. 295 pp. [Available from Andrew Isles Natural History Books]

Bailey, Col. (2016). Lure of the Thylacine: True Stories and Legendary Tales of the Tasmanian Tiger. Echo Publishing. 227 pp. [Available from Amazon]

Barbeliuk, Anne “Ethical issues may halt tiger cloning,”. The Mercury 9211999.

Barbeliuk, Anne. (1999, September 9). A tiger by the tail. The Mercury, Hobart, p. 19.

Barbeliuk, Anne. (1999, September 10). Thylacine cloning hit as mission impossible. The Mercury, p. 9.

Barbeliuk, Anne. (2001, October 30). Expert’s fears as vote favors tiger cloning. The Mercury, Hobart, p. 5.

Barrow, M. V. (2012). Framing the Thylacine. Society and Animals 20(1): 109-110.

Bartholomai, A. (1977). The fossil vertebrate fauna from Pleistocene deposits at Cement Mills, Gove, southeastern Quuensland. Mem. Qld. Mus. 18: 69-73.

Bartosch, Roman. (2016). Ghostly Presences: Tracing the Animal in Julia Leigh’s The Hunter, pp. 259-275. In: Herman, David (ed.). Creatural Fictions (Palgrave Studies in Animals and Literature series). US: Palgrave Macmillan US. [Abstract]

Baudement, G. (1849). Thylacine. In: D'Orbigny, M. C. (ed.). Dictionnaire Universel D'Histoire Naturelle, Vol. 12. Paris: Renard, Martinet et Cie.

Baynes, Alexander and Jones, Barbara. (1993). The mammals of Cape Range peninsula, Western Australia, pp. 207-226. In: Humphreys, W. F. (ed.). The Biogeography of Cape Range, Western Australia. Perth: Western Australian Museum.

Baynes, Alexander, Merrilees, D. and Porter, J. K. (1975). Mammal remains from the upper level of a late Pleistocene deposit in Devil's Lair, Western Australia. J. Roy. Soc. W. Aust. 49: 88-90.

BBC Online “Tasmanian tiger may growl again,”. May 14, 1999,

Beale, Bob. (2005). Bogus focus. The Bulletin with Newsweek 123(6464): 24-25. ["The image of a thylacine snaring a chicken in 1921 is one of the best-known photographs of the extinct species and was widely published in newspapers, books, magazines, and documentary films. However, a Tasmanian University researcher suggests that the famous 1921 snap is staged."]

Beaton, J. M. (1991). Cathedral Cave: a rockshelter in Carnarvon Gorge, Queensland. Queensland Archaeological Research 8: 33-84. [relevant citation?]

Beck, R. M. D. (2008). A dated phylogeny of marsupials using a molecular supermatrix and multiple fossil constraints. Journal of Mammalogy 89: 175-189.

Beddard, Frank E. (1891). On the pouch and brain of the male Thylacine. Proceedings of the Zoological Society of London 59(1): 138-145[-148?].

Beddard, Frank E. (1903). Exhibition of and remarks upon sections of the ovary of the thylacine. Proceedings of the Zoological Society of London 1903: 116.

Bell, E. A. (1965). Mrs Roberts and the Tasmanian tiger, pp. 103-110. In: (editor?). An historic centenary. Roberts, Stewart S Co. Ltd. 1865–1965. Hobart: Fuller's Book Shop. [According to Calaby: "History of a firm of wool brokers. Ch. 14 contains some interesting historical information and a photograph of a captive group of an adult female and three subadult thylacines. Mrs Roberts had a well-known private zoo"]

Bell, E. A. (1967). “Tigers” were her hobby. Australian Women’s Weekly, 10/5/1967, pp 12-13.

Bell, E. A. (1975). Thylacine. Archives Office of Tasmania.

Bell, E. A., ms paper NS 463/2, Archives Office of Tasmania.

Bell, E. A., Thylacine Reports--Queen Victoria Museum--Launceston, unpublished ms, Archives Office of Tasmania NS 896/1-39.

Bennett, David. (2003). Often crude and quaint: some Australian conceptions of nature, ecology, and rock-art. Before Farming 2003(4): 1-11. [Abstract]

Bensley, B. A. (1903). On the evolution of the Australian Marsupialia; with remarks on the relationships of the marsupials in general. Trans. Linn. Soc. Lond. 9: 83-218.

Benson, S. (1999a, September 8). Tiger cloning project begins. The Mercury, Hobart, p. 7.

Benson S (1999b) Test tube tiger: extinct thylacine to be cloned. The Daily Telegraph, 8 September

Benson, S. (2000, May 5). Tasmanian tiger may live again. The Daily Telegraph, p. 6.

Benson, S. (2001, March 28). Tickling tiger to life: Thylacine clone bid. The Mercury, p. 36.

Beresford, Quentin. (1985). Tradegy of the Tasmanian tiger. The Islander 1985: 12-14.

Beresford, Quentin and Bailey, Gary. (1981). Search for the Tasmanian Tiger. Hobart, Tasmania: Blubber Head Press. 54 pp.

Berns, Gregory S. and Ashwell, Ken W. S. (2017). Reconstruction of the Cortical Maps of the Tasmanian Tiger and Comparison to the Tasmanian Devil. PLoS ONE 12(1): e0168993.

Binks, C.J. 1980. Explorers of Western Tasmania. Launceston: Mary Fisher Bookshop.

Blackwell, W. 1951. Thylacines: Typed interview notes by Linsay Crawford, 27/11/1951. Thylacine Expeditionary Research Team Files, Queen Victoria Museum, Launceston.

Boardman, F. (1945). Some points on the external morphology of pouch young of the Marsupial, Thylacinus cynocephalus Harris. Proceedings of the Linnean Society of New South Wales 70: 1-8.

Boroughs, D. (2000). Stripes and shadows. Timbila: Magazine of the South African National Parks, 2 (1), np. [relevant citation?]

Bowdler, Sandra. (1984). Hunter Hill, Hunter Island. Terra Australis 8: xv + 148 pp. [a single thylacine molar tooth was found, dated to roughly 15,400 BP; still the only known record of the species from the Bass Strait islands as far as I am aware]

Boyce, (Peter) James. (2006). An environmental history of British settlement in Van Diemen's Land: The making of a distinct people, 1798-1831. Doctoral dissertation, University of Tasmania.

Boyce, (Peter) James. (2008). Return to Eden: Van Diemen's Land and the early British settlement of Australia. Environment and History 14(2): 289-307. [Europeans dogs much faster than thylacines; thylacines seem to have retreated because of the dog to less open country]

Brandl, E. J. (1972). Thylacine designs in Arnhem Land rock paintings. Archaeology and Physical Anthopology in Oceania 7(1): 24-30.

BRAUN, M. J. (1992). Phylogenetic relationships of the thylacine (Mammalia: Thylacinidae) among dasyuroid. In Proc. R. Soc. Lond. B (Vol. 250, pp. 19-27).

Brehm, Alfred Edmund. (1895). Tierleben (Life of Animals). Chicago, Illinois: A. N. Marquis & Co. Publishers.

Breton, W. H. (1847). Description of a large specimen of Thylacinus harrisii Exhibit of a large specimen of Thylacinus Harrisii. Tasmanian Journal 3: 125-126.

Bristow, Mila. (2012) Tracking down rock art records of thylacines and sailing ships. Australian Forest Grower 35(3): 19. [Abstract]

Brogden, Stanley. (1948). Tasmanian Journey. Melbourne: Morris & Walker for Pioneer Tours.

Brook, S. (2000, May 5). Thylacine clone no longer just a paper tiger. The Australian, p. 1.

Broom, R. (1896). Report on a bone brec[c]ia near Wombeyan Caves, N. S. W.. Proc. Linn. Soc. N. S. W. 21: 48-61.

Brown, Robert ("Bob"). (1972). The Tasmanian tiger. Skyline 1972: 29-31.

Brown, Robert ("Bob"). (1973). Has the thylacine really vanished? Animals 15(9): 416-419.

Brown, Robert ("Bob"). (1983). Has the last thylacine gone to ground there hope for our tiger? Tasmanian Mail, 16 August, p. 8.

Bryant, Sally L. and Jackson, J. (1999). Tasmania's Threatened Fauna Handbook: What, Where and How to Protect Tasmania's Threatened Animals. Threatened Species Unit, DPIWE, Hobart

Bryant, Sally and Squires, Tim. (2009). Animals of Tasmania: Wildlife of an Incredible Island. Quintus Publishing. 80 pp.

Bulte, E. H., Horan, R. D. and Shogren, J. F. (2003). Is the Tasmanian Tiger extinct? A biological-economics re-evaluation. Ecological Economics 45(2): 273-281. [Abstract]

Buk, Steve. (1985). Just how extinct is Tasmania's tiger? International Wildlife 15(4): 36-39.

Bulmer, S. (1964). Radio Carbon dates from New Guinea. Journal of the Polynesian Society 73(3): 327-328. [relevant citation?]

Bulmer, S. E. (1966/76). The Prehistory of the New Guinea Highlands. Unpublished MA thesis, Auckland: University of Auckland. Microfiche edition produced 1977 with new introduction.

Bulmer, S. (1974). Working paper in Anthropology & Archaeology. Linguistics and Maori Studies No. 30, Archaeology. Unpublished Report, Department of Anthropology, University of Auckland.

Bulmer, Susan. (1979). Archaeological Evidence of Prehistoric Faunal Change in Highland Papua New Guinea. Unpublished paper to Australian and New Zealand Association for the Advancement of Science Congress, Section 25A. Auckland.

Bunk, S. (1985). Just how extinct is Tasmania's tiger. International Wildlife 15(4): 36-39.

Burbury, Alfred. (2000). Alfred Burbury Memories from the Chronicles of the Burbury Family. Oatlands District Historical Society Chronicle 1: 29. [ "when the dogs had gone [during the second half of the nineteenth century], native tigers took over, notably in the east and around Tooms Lake."]

Burbury, F. (1953). Letter to Eric Guiler, 1953.

Burrell, Harry. (1921). The Tasmanian Tiger or Wolf (photo). Australian Museum Magazine 1(3): 62.

Buttrose, Larry. (1987). Yowies and Thylacines Alive: One Man's Vision. The Living Australia Magazine 1987(May): [pagination?].

Calaby, J. H. (1983). Thylacine, 10: 52-53. Australian Encyclopaedia, 4th ed.

Calaby, J. H. and White, C. (1967). The Tasmanian Devil (Sarcophilus harrisii) in northern Australia in recent times. Aust. J. Sci. 49: 473-475. [relevant to Thylacinus?]

Cambrian. 1855. Notes on the natural history of Australasia, letter third. Melbourne Monthly Magazine 1 (October): 360-362.

Carlos, Colin J., Bond, Alexander L. and Burgio, Kevin R. (Preprint, 2017). Estimating the extinction date of the thylacine accounting for unconfirmed sightings. bioRxiv 123331. doi:

Case, Judd A. (1985). Differences in prey utilisation by Pleistocene marsupial carnivores, Thylacoleo carnifex (Thylacoleonidae) and Thylacinus cynocephalus (Thylacinidae). Australian Journal of Mammalogy 8(1-2): 45-52.

Chaloupka, George. (1975). Fallen emblem – or lingering star? EZ Review (1):2-4. [possible rock art depcitions of Thylacinus]

Chaloupka, George. (1977). Aspects of the chronology and schematisation of two prehistoric sites on the Arnhem Land Plateau. In P.J. Ucko (ed.), Form in indigenous art: schematisation in the art of Aboriginal Australia and prehistoric Europe, pp. 243-59. Australian Institute of Aboriginal Studies, Canberra.

Chaloupka, George. (1993). Journey in Time: The World’s Longest Continuing Art Tradition. Chatswood, NSW: Reed. [possible rock art depcitions of Thylacinus]

Churcher, C.S. (1985). Dental functional morphology in the marsupial sabre-tooth Thylacosmilus atrox (Thylacosmilidae) compared to that of felid sabre-tooths. Australian Mammalogy 8: 201–220. [J. H. Calaby: "[has] information on Thylacinus"]

Cininas, Jazmina. (2008). Antipodean Bestiary: Reconstructions of Native Fauna and National Identity in the Work of Eleven Contemporary Australian Artists. PAN: Philosophy Activism Nature 5: 21-39. [automatic download]

Clark, Tim W.; Reading, Richard P.; Wallace, Richard L.; Wilson, Barbara A. (2002). If the Tasmanian Tiger Were Found, What Should We Do? An Interdisciplinary Guide to Endangered Species Recovery. Endangered Species Update 19(4): 194-200.

Claude, C. (1996). Der Beutelwolf. Thylacinus cynocephalus Harris, 1808. Leben und Sterben einer Tierart. Zurich: Zoologisches Museum der Universität Zürich.

Clegg, J., 1978. Pictures of striped animals: which ones are thylacines? Archaeology and Physical Anthropology in Oceania 13(1): 19-29.

Clucas, E. (1978). Tracing the Decline of the Thylacine. Tasmaniana Library.

Cochrane, Peter. (undated). Bizarria: Tasmanian Tiger - Vampire Dog? Available at:

Colbron-Pearse, D. (1968). Tame tiger. Walkabout 1968(December): 8.

Coleman, R. (1976). There's a strange, strange beastie out there. Melbourne Herald, 20 March, p. 29.

Colgan, Don et al. (2000). The Thylacine Project. Australasian Science 21: 1. [same publication as (Colgan & Archer, 2000)?]

Colgan, D. (2002). Thylacine cloning is a long shot. Australasian Science, 23(6), 15-15.

Colgan, Don and Archer, Michael. (2000). The Thylacine Project. Australasian Science 21(1): 21. ["Don Colgan and Mike Archer respond to criticisms of plans to clone the extinct Tasmanian tiger."]

Collins, L. R. (1973). Monotremes and marsupials. A reference for zoological institutions. Washington D.C.: Smithsonian Institute Press. v + 323 pp.

Colliver, F. S. (1938). Cave hunting in Victoria. Vict. Nat. 54: 152-155.

Cook, D. L. (1963). Thylacinus and Sarcophilus from the Nullabor Plain. Western Australian Naturalist 9(2): 47-48. [Thylacine molar found in Tasmanian devil coprolite]

Corbett, L. K. (1995). The Dingo in Australia and Asia. Sydney: University of NSW Press.

Corbett, L. K. (2001). The Dingo in Australia and Asia. Adelaide: J. B. Books.

"Correspondent". (1924). ["article on the killing of a tiger at Waratah by C. Penny" according to (Guiler & Godard, 1998:242)]. Weekly Courier, 17 January, pp. 26-46.

Cosgrove, Richard. (1984). Aboriginal Economy and Settlement in the Central Highlands, Tasmania.Hobart: National Parks and Wildlife Service. Occasional Paper 8.

Cosgrove, Richard and Allen, Jim. (2001). Prey Choice and Hunting Strategies in the Late Pleistocene: Evidence from Southwest Tasmania, pp. 397-430. In: Anderson, A., O'Connor, S. and Lilley, I. (eds.). Histories of Old Ages: Essays in Honour of Rhys Jones. Canberra: Coombs Academic Publishing.

Cosgrove, Richard, Field, Judith, Garvey, Jillian, Brenner-Coltrain, Joan, Goede, Albert, Charles, Bethan, Wroe, Steve, Pike-Tay, Anne, Grün, Rainer, Aubert, Maxime, Lees, Wendy and O'Connell, James. (2010). Overdone overkill – the archaeological perspective on Tasmanian megafaunal extinctions. Journal of Archaeological Science 37: 2486-2503.

Crawford, L. (1951). "Thylacines: Typed interview notes by Linsay Crawford", 27 November 1951. Thylacine Expeditionary Research Team Files, Queen Victoria Museum, Launceston.

Crawford, W. (2002, June 1). Tasmanian pup dream. The Mercury, Hobart, p. 25.

Crawford, L. D. (1952). Tasmanian Tiger. Launceston: Queen Victoria Museum.

Crisp, E. (1855). On some points relating to the anatomy of the Tasmanian Wolf (Thylacinus) and of the Cape Hunting Dog (Lycaon pictus). Proceedings of the Zoological Society of London 1855: 188-191.

Crook, Bronte and Hiraki, Arisa. (2016). An Examination of Feeding, Digestion, and Cellular Metabolism in the Tasmanian Tiger, Thylacinus cynocephalus. Zool 430. [previous version]

Cunningham, Daniel John. (1877). On the Myology of the Shoulder and Upper Arm Thylacine, Cuscus, and Phascogale [4 pp.; exact pagination unknown]. In: Report of the British Association for the Advancement of Science, 47th. Meeting, Plymouth, 1877. London: John Murray.

Cunningham, Daniel John. (1878a). The Nerves of the Fore-Limb of the Thylacine (Thylacinus Cynocephalus or Harrisii) and Cuscus (Phalangista Maculata). J. Anat. Physiol. 12(Pt 3): 427-433.

Cunningham, Daniel John. (1878b). The Intrinsic Muscles of the Hand of the Thylacine (Thylacinus Cynocephalus), Cuscus (Phalangista Maculata), and Phascogale (Phascogale Calura). J. Anat. Physiol. 12(Pt 3): 434–444.

Cunningham, Daniel John. (1881a). The Nerves of the Hind-Limb of the Thylacine (Thylacinus Harrisii or Cynocephalus) and Cuscus (Phalangista maculata). J. Anat. Physiol. 15(Pt 2): 265–277.

Cunningham, Daniel John. (1881). Report on some points of anatomy of the thylacine (Thylacinus cynocephalus), cuscus (Phalanger maculata) and phascogale (Phascogale calura) collected during the voyage of H.M.S. Challenger in the years 1873-1876: with an account of the intrinsic muscles and nerves of the mammalian pes. Report on the Marsupialia. Voyage of the H.M.S. Challenger, Zoology 5(16): 1-192. Edinburgh: Printed by Neill and Co. for Her Majesty's Stationary Office. [includes 13 plates] [Available from Andrew Isles NHB]

Czechura, G. V. (1984). Modern sightings of the thylacine - what do they tell us? Skeptic 4(4): 1-3,6.

Davis, William E. Jnr. (2007). Tasmania: A Natural History. Chipping Norton, NSW: Surrey Beatty & Sons. 269 pp.

Dawson, L. (1982). Taxonomic status of fossil thylacines (Thylacinus, Thylacinidae, Marsupialia) from late Quaternary deposits in eastern Australia, pp 517-525. In: Archer, Michael (ed.). Carnivorous Marsupials. Mosman, N.S.W.: Royal Zoological Society of New South Wales.

Dawson, L. (1985). Marsupial fossils from Wellington Caves, New South Wales; the historic and scientific significance of the collections in the Australia Museum, Sydney. Records of the Australian Museum 37: 55-69.

Dawson, L. and Augee, M. L. (1997). The late Quaternary sediments and fossil cave vertebrate fauna from Cathedral Cave, Wellington Caves, New South Wales. Proc. Linn. Soc. N.S.W. 117: 51-78.

Day, David. (1981). The Doomsday Book of Animals: A Natural History of Vanished Species. New York, N.Y.: The Viking Press.

Dayton, L. (2002, May 29). Tiger on comeback trail. The Australian, p. 1.

de Groen, Frances. The thylacine hunter [online]. Quadrant, Vol. 24, No. 6, June 1980: 67.

de Moeller, O. (1998). On the trail of a Thylacine. West Australian, 5 January 1998, p. 7.

De Vis, Charles W. [In. Anon.] (1893). A thylacine of the earlier nototherian period in Queensland. Proceedings of the Linnean Society of New South Wales (Abstract) 29th November, 1893. [Republished 1894, Zool. Anz. 17 : 47.]

De Vis, Charles W. (1894). A thylacine of the earlier nototherian period in Queensland. Proceedings of the Linnean Society of New South Wales (second series) 8: 443-447.

De Vos, Ricardo. (2007). Extinction Stories: Performing Absence(s), pp. 183-195: In: Simmons, L. and Armstrong, P. (eds.). Knowing Animals. Leiden: Brill.

Dixon, Joan M. (1989). Thylacinidae (Chapter 20), pp. 549–559. In: Walton, D. W. and Richardson, B. J. (eds.). Fauna of Australia Mammalia. Canberra: Australian Government Publishing Service.

Dixon, Joan M. (1991). The Thylacine – Tasmania's tiger. Melbourne, Victoria: Museum of Victoria. 9 pp. ill., map.

Doherty, K. (1977). When we caught a tiger. In: N.W. Tasmania Short Stories and Articles. Boat Harbour: Tasmanian Fellowship of Australian Writers, North West Branch.

Dortch, J. and Wright, R. 2010. Identifying paleoenvironments and changes in Aboriginalsubsistence from dual-patterned faunal assemblages, south-western Australia. Journal of Archae-ological Science 37: 1053-1164. [relevant citation?]

Douglas, Athol M. (1986a). Tigers in Western Australia? New Scientist, 24 April, pp. 44-47.

Douglas, Athol M. (1986b). Thylacine lives? (letter to the editor). New Scientist, 24 July, 111(1518): 63.

Douglas, Athol M. (1990). The Thylacine: A Case for Current Existence on Mainland Australia. Cryptozoology/Cryptozoologie 9: 13-25.

Dunn, Ashley and Colgan, Don. (2002). [Collection of two articles on a project to use DNA from the thylacine to bring it back from extinction]. Australasian Science 23(6): 14-15.

Dwyer, P. D., 1982. Wildlife conservation and tradition in the highlands of Papua New Guinea. Pp. 173-89 in Traditional Conservation in Papua New Guinea: Implications for Today ed by L. Morauta, J. Pernetta and W. Heaney. Institute of Apptied Social and Economic Research, Boroko, Papua New Guinea. [relevant citation?]

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Freeman, Carol J. (2004c). ‘Figuring Extinction: The First Illustration of a Thylacine (Tasmanian tiger) in a Scientific Work’, Cross Fertilisations: Literature, Science and Nature. An Interdisciplinary Conference July 16-18 2004 Programme, July 2004, Chichester, UK, pp. 3-4. (2004) [Conference Extract]

Freeman, Carol J. (2005a). Is this picture worth a thousand words? An analysis of Harry Burrell's photograph of a thylacine with a chicken. Australian Zoologist 33: 1-16.

Freeman, Carol J. (2005b). Figuring extinction: Visualizing the thylacine in zoological and natural history works 1808-1936. Unpublished doctoral dissertation. University of Tasmania: Hobart, Australia.

Freeman, Carol J. (2005c). The Abject Thylacine: interactions between the real and the representation. Animals & Society: Inaugral Conference of the Animals & Society (Australia) Study Group, July 2005, Crawley, WA, pp. 36-36. [Conference Extract]

Freeman, Carol J. (2005). From Tasmania to Knowsley: John Gould's Thylacines. Stanley Estates Newsletter, Earl of Derby, Liverpool, 17, August. [Internal Newsletter]

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Roberts, M.G. 1912. Diary, 1/7/1911 – 26/12/1912. Roberts collection, State Archives Office, Hobart.

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Roberts, M.G. 1919. Letter to H.A. Longman, 27/3/1919. Archives collection, Queensland Museum, Brisbane.

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Robovský, Jan, Sleightholme, Stephen R., Vohralík, Vladimír and Benda, Petr. (2015). Specimens of Thylacinus cynocephalus in collections of the Czech Republic (Mammalia: Thylacinidae). Journal of the National Museum (Prague), Natural History Series 184(2): 43-50.

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Roux, J. (1772). Journal of the voyage made on the King's ship, the Mascarin, commanded by M. Marion Knight of the Royal and Military Order of St Louis Fireship Captain accompanied by the flute, the Marquis de Castries, commissioned to make a voyage to the island of Tahiti or Cythea and to expore the Southern Lands, thence proceeding to New Holland, to New Zealand etc .etc. (Maryse Duyker, Trans.). In E. Dyker (Ed.), The Discovery of Tasmania: journal extracts from the expeditions of Abel Janszoon Tasman and Marc-Joseph Marion Dufresne 1642 and 1772 (1992 ed., pp. 38-43). Hobart: St David's Park Publishing. [first known reported thylacine sighting by Europeans: “We have not seen any quadrupeds other than a little tiger [qu’un petit Tigre] which ran away
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West, [of Burnie]. ca1970. Handwritten Interview Notes. Thylacine Expeditionary Research Team Files, Queen Victoria Museum, Launceston.

West, John. (1852). The History of Tasmania, Volume 1. Launceston: Henry Dowling. 336 pp.

Whelan, Barbara. (2007). The extinction of the Thylacine (Thylacinus cynocephalus): Quantifying the effects of competition, hunting, disease and habitat alteration. Thesis.

Whitely, G. P. (1973). I remember the thylacine. Koolewong 2(4): 10-12. [p. 10-11?]

Whitley, G.P. 1975. Thylacine Folder. Australian Museum Archives, Sydney.

Whitticker, Mark. (1997). Look! There's one. -Tasmanian tigers aren't extinct, they're everywhere say believers; but scientists say thylacine hunters are just chasing their tails. Australian. Magazine, 15-16 Nov 1997, p.12-18.

Wild, Mary. (1997). Tigers in the outback. E: The Environmental Magazine 8(3): 22. [wordcount: 405; unfortunately contains several factual errors]

Wilford, J. N. (1980). A new search for the rare tiger. Intern. Herald Tribute, 6 June.

Williams, Dominic L. G. (1980). Catalogue of Pleistocene vertebrate fossils and sites in South Australia. Proc. R. Soc. S. Aust. 104: 101-115. [relevant citation?]

Williams, Louise. (1997). Tassie tiger sighting claim in Irian Jaya. The Sydney Morning Herald, 15 April.

Williams, Michael "Mike" and Lang, Rebecca "Ruby". (2010). Australian Big Cats: An Unnatural History of Panthers. Hazelbrook, NSW: Strange Nation Publishing.

Wilson, A. ca1922w. Notes on the Tiger. Unpublished manuscript, Archie Wilson collection, Tasmanian Library, State Library of Tasmania, Hobart.

Woodburne, M. O. (1967). The Alcoota Fauna, central Australia. Bulletin of the Bureau of Mineral Resources Geology and Geophysics, Australia 87: 1-187.

Woodford, James. (1995). New bush sighting puts tiger hunter back in business. The Sydney Morning Herald, 30 January.

Woodford, J. (1999). Unleashing the thylacine. The Age, 1.

Woodford, James. (2000, May 5). Get a life, scientists tell extinct tiger. Sydney Morning Herald, Late ed. p. 3.

Woodford, James. (2001, November 24–25). Extinction for the time being. Sydney Morning Herald, Weekend Edition, Spectrum, 4-5.

Woods, A. (1977). Tiger, tiger in the night. Sydney Morning Herald, 25 August, p. 7.

Wright, Bruce J. (1972). Rock engravings of striped marsupials: the Pilbara Region, Western Australia. Archaeology and Physical
Anthropology in Oceania 7(1): 15-23. [Abstract]

Wroe, Stephen. (2001). Maximucinus muirheadae, gen. et sp. nov. (Thylacinidae: Marsupialia), from the Miocene of Riversleigh, north–western Queensland, with estimates of body weights for fossil thylacinids. Australian Journal of Zoology 49: 603-614.

Wroe, Stephen and Attard, Marie. (2012). The thylacine myth. Australasian Science 33(5): 19-22.

Wroe, S., Clausen, P., McHenry, C., & Moreno, K. (2007). Finite element modeling of feeding behavior in the thylacine and wolf: A novel test for convergence. In JOURNAL OF VERTEBRATE PALEONTOLOGY (Vol. 27, No. 3, pp. 169A-169A).

Wroe, Stephen; Clausen, Philip; McHenry, Colin; Moreno, Karen and Cunningham, Eleanor. (2007). Computer simulation of feeding behaviour in the thylacine and dingo as a novel test for convergence and niche overlap. Proc. R. Soc. B 274: 2819–2828.

Wroe, Stephen, Lowry, Michael B. and Anton, Mauricio. (2008). How to build a mammalian super-predator. Zoology 111: 196-203.

Wroe, Stephen, McHenry, Colin and Thomason, Jeffrey. (2005). Bite club: comparative bite force in big biting mammals and the prediction of predatory behaviour in fossil taxa. Proc. R. Soc. B 272(1563): 619-625. [Also available from Royal Society Proceedings B] [Supplementary data]

Wroe, Stephen and Milne, Nicholas. (2007). Convergence and remarkably consistent constraint in the evolution of carnivore skull shape. Evolution 61(5): 1251-1260.

Wroe, Stephen and Musser, A. (2001). The skull of Nimbacinus dicksoni (Thylacinidae: Marsupialia). Australian Journal of Zoology 49(5): 487-514. [Abstract]

Yates, Adam M. (2015). Thylacinus (Marsupialia: Thylacinidae) from the Mio-Pliocene boundary and the diversity of Late Neogene thylacinids in Australia. PeerJ 3: e931.

Yendall, D. (1982). Search for the thylacine. Wildlife (International) 24(May): 182-183.

Scientific Papers
Royal Society of Tasmania’, Courier, 16 June 1858, p.2; or ‘Town talk and table chat’, Cornwall Chronicle, 27 March 1867, p.4.

[Theophilus Jones], ‘Through Tasmania: no.35’, Mercury, 26 April 1884, supplement, p.1.

‘Parliament’, Launceston Examiner, 1 October 1886, p.3.

Tasmanian Mail, 3 September 1887

E Richall Richardson, ‘Tour through Tasmania: no.89: cattle branding’, Tasmanian Tribune, 14 March 1878, p.3.

‘Buckland’, Mercury, 15 August 1884, p.3.

Theophilus Jones, ‘Through Tasmania: no.61’, Mercury, 8 November 1884, supplement, p.1.

Theophilus Jones, ‘Through Tasmania: no.49’, Mercury, 26 July 1884, p.2.

Theophilus Jones, ‘Through Tasmania: no.35’, Mercury, 26 April 1884, supplement, p.1.

Theophilus Jones, ‘Through Tasmania: no.56’, Mercury, 20 September 1884, p.2.

Theophilus Jones, ‘Through Tasmania: no.35’, Mercury, 26 April 1884, supplement, p.1.

‘House of Assembly’, Mercury, 24 October 1884, p.3.

‘Meeting at Buckland’, Mercury, 14 August 1884, p.2; ‘The native tiger’, Mercury, 25 September 1885, p.2.

Editorial, Mercury, 28 March 1884, p.2.

Popular articles
Psychic Australian November 1976 [author? volume? pagination?]

Psychic Australian January 1977 [author? volume? pagination?]

Paranormal and Psychic Australian October 1977 [author? volume? pagination?]

Paranormal and Psychic Australian May 1978 [author? volume? pagination?]

Downes, Jonathan (ed.). (year?). Animals & Men - The Journal of the Centre for Fortean Zoology, Issue 19 [contains an article called "A British Thylacine?"]

The Imperial Natural History Picture Book New York & London: George Routledge and Sons, 1886, 64 pp.

Explorer's Journal, June 1980, USA: The Explorer's Club. ["Devils, Quolis, Dunnarts and Thylacines - a survey of Tasmanian mammals"]

Nature, Temporality and Environmental Management: Scandinavian and Australia Perspectives on Peoples and Landscapes. [p. 50-54]

Boogeymen. (aired 16 November 2015). Season 2, Episode 9: Beast of Buderim. "A strange creature believed to be related to the Tasmanian Tiger is spotlighted". source:

Mohr, M. (Producer). (2000, May 4). Bringing the Tasmanian tiger back to life. In ABC Online: 7.30 Report.

O‘Neill, P. (Producer/Director/Writer). (2002). End of extinction: Cloning of the Tasmanian tiger. Silver Spring, MD: Discovery Channel/TLC.

Wellington, C. (1996). The Tasmanian Tiger. The Definitive Documentary, (videotape). Hobart: Winning Post Productions.

Beyond the Jaws of Extinction, Chris Packham (host), originally aired 30 September 1998,

Other Multimedia:

The tragedy and myth of the Tasmanian Tiger. (2001). CD-ROM. Hobart, Tasmania: Roar Film & Screen Tasmania. ["Tells the story of the Tasmanian Tiger by drawing on primary source material and using science to understand the species and its behaviour." (source)]

Tasmanian tiger: Thylacinus cynocephalus: alive & well : new taped interviews of genuine sightings since 1980. (2004). Terry, Edward "Ned" Vincent. CD Audiobook. Dairy Plains, Tasmania: Self Published. [alternative title: "Tasmanian Tiger: Alive & Well"; published in 1999?]

Save the Tassie tiger! (2000). Developed by R3 Interactive. Adelaide, South Australia: EcoTigers Support Group. + 1 booklet (18 pp.).

Phenomena Magazine. Secrets of Science - The Resurrection of the Tasmanian Tiger (Documentary, F 2005)

Websites [Murray McAllister's website] [research project] [Some of Col Bailey's "Tiger Tales" newspaper column articles] [photograph of Elias Churchill's hut, capturer of the last known wild thylacine in the Florentine Valley in 1933]

"In June 2009, the Cranbourne Journal (predecessor of the Casey Weekly) ran an article on a possible sighting of a Tasmanian tiger crossing Chevron Avenue, Cranbourne South, in 2001. Another Cranbourne resident claimed to have seen a tiger in Tooradin in 2000." (source) (Ubirr rock art site contains depiction of thylacine)

Fictional Accounts
NB: this section includes non-fiction reviews about fictional works.

Aronson, Linda. (year?). Naturally Rude. Puffin. [incomplete citation]

Atwood, M. Thylacine Ragout.” 2006. The Tent, 73-75. [relevant citation?]

Barzilai, Shuli. 2008. “Unfabulating a Fable, or Two Readings of ‘Thylacine Ragout.’” In Once upon a Time: Myth, Fairy Tales and Legends in Margaret Atwood’s Writings, ed. Sarah A. Appleton. Cambridge: Cambridge Scholars Publishing, pp. 127–50. [relevant citation?]

Black, Tony. (Due 1 October 2014). The Last Tiger. Cargo Publishing. 324 pp. [Available from Amazon] (

Campbell, Justine and Sarah Hamilton. (2013). They Saw a Thylacine. Currency Press. 66 pp. [Kindle edition] [a play]

Carey-Wells, Penny and Perndt, Diane. (2006). Pockets & corners : furry facts and thylacine fiction in the heartlands of Tasmania. Kingston Beach, Tasmania: Cloud Design. 109 pp.

Carr, Aaron. (2016). Tasmanian Tiger. New York, New York: AV² by Weigl.

Darlison, Aleesah (text) and McGrath, Shane (illustrator). (2016). Stripes in the Forest: The Story of the Last Wild Thylacine. Big Sky Publishing. 32 pp.

Dying Breed. Dir. Jody Dwyer. Ambience Entertainment, 2008.

Chauncy, Nan. (1957). Tiger in the Bush. London: Oxford University Press. 172 pp.

Crew, G. and Wilson, M. (2003). I saw nothing: the extinction of the thylacine. Victoria: Lothian Books.

Cromer, William "Bill". (2003). Thylacine conspiracy: Intrigue and Suspense in Tasmania. Springboro, Ohio: Just my Best Publishing. 246 pp. [Bill Cromer: "Reprinted 2003 in Tasmania with a different dust cover" (260 pp.?)]

D'Ath, Justin. (1996). Infamous. [Publisher?] ISBN: 9781875930036.

de Groen, Frances. (1980). The thylacine hunter. Quadrant 24(6): 67. [a poem]

Domico, Terry. (2005). The Last Thylacine. Washington & Strathfield, N. S. W: Turtleback Books, 232 pp. [Google Books Preview]

Evans, Gareth. (2005). The Thylacine Conspiracy. E & F Publications. 256 pp.

Ford, Catt and Kennedy, Sean. (2009). Dash and Dingo. Dreamspinner Press, 310 pp.

Frauca, Harry. (1969). Striped Wolf: A Bush Adventure. Melbourne: Heinemann. 109 pp.

Freeman, Carol. (2013). The last image: Julia Leigh's The hunter as film, [pagination?]. In: Johnston, Jay and Probyn-Rapsey, Fiona (eds.). Animal Death. Sydney University Press. 348 pp.

Gambles, Harriet and Gambles, Patrick. (2001). Tiger Trail - A Tasmanian Epic. Tasmania: Regal Publications.

Gordon, Victoria. (2010). Wolf in Tiger's Stripes. Gale, 252 pp. [Google Books]

Hardy, Maurice Athol. (2012). Queen Mallaegania and the Valley of the Demon: Plus Night of the Thylacine. Createspace. 168 pp. [relevant citation?]

Hartnett, Sonya. (1995). Sleeping Dogs. Viking Juvenile, 144 pp. [Available from Amazon]

Hartnett, Sonya. (2007). Stripes of the Sidestep Wolf. Candlewick Press, 208 pp.

The Hunter. Dir. Daniel Nettheim. Porchlight Films, 2011.

Irby, Kenneth Alan (lyrics) and Cameron, Rod (music). (1996). Our Tassie tiger song. Wynyard, Tasmania: Self published. 3 pp. ill.

Isham, Marion and Isham, Steve. (1996). Tasmanian Tiger. Bandicoot Books. 30 pp.

Jenyns, L, Tasmanian Tiger: The Mystery of the Thylacine, Tasmanian Museum and Art Gallery, Hobart, pp. Broadsheet (1998) [Recorded Creative Work]

Jonsberg, Barry. (2011). Blacky Blasts Back: On the Tail of the Tassie Tiger. Allen & Unwin.

Leigh, Julia. (1999). The Hunter. London: Faber and Faber / New York: Penguin.

Martin, John (author) and Connors, Maria (illustrator). Who Knew Tasmanian Tigers Eat Apples! (Windy Mountain). Privately published? 288 pp.

Maynard, Leigh. (2004). How the Tasmanian Tiger got its Stripes. Scholastic. 32 pp.

McMillan, Dawn. (2013). Red Riding Hood and the Tassie Tiger. Publisher?

NcNair, Leonie. (1991). Tyler the Tasmanian Tiger. Launceston, Tasmania: Holiday Link. 28 pp.

Nowra, Louis. (2012). Into That Forest. Crows Nest, N.S.W.: Allen &​ Unwin. 172 pp.

Ordinans, Nicholas J. (2013). Where There Be Tygers. Self Published. 281 pp. [Available from Amazon]

Osborne, Lynda. (2015). Status: Presumed Extinct. Pegasus Pony. 260 pp. [Kindle edition]

Reed, Bill. (1982). Ihe. Hyland House.

Reed, Bill. (2015). Ihe. Dandenong, Victoria: Reed Independent.

Roberts, Beth. (1990). The Magic Waterfall. Kingston, Tasmania: Rainbow Books. 44 pp.

Rose, Heather. (1999). White Heart. Anchor Publishing, 312 pp.

Salmon, Michael. (1986). The Great Tasmanian Tiger Hunt. Templestowe, Victoria: Lamont Books. 28 pp.

Saxby, M. (2011). The Dream of the Thylacine by Margaret Wild and Ron Brooks. Magpies -Victoria Park 26(1): 10-11.

Small, Mary. (1985). Grandfather's Tiger: Search for the Tassie Tiger. North Ryde, New South Wales: Methuen. 94 pp.

Small, Mary. (1989). Grandfather's Tiger: Search for the Tassie Tiger. Sydney, New South Wales: Collins. 104 pp.

Tuckey, Noel G. (2008). Thylacine Man. Glen Waverley, Vic: Sid Harta. 354 pp.

Tully The Tasmanian Tiger And Friends. No 2. Colouring-In and Activity Book. 64 pp.

Van Essen, Susanna. (1996). Denying extinction?: the thylacine in children's literature. Reading Time 40(3): 12-15.

Wild, Margaret and Brooks, Ron. (2011). The Dream of the Thylacine. Crows Nest, N.S.W.: Allen &​ Unwin [Children's?].

Wild, Margaret and Brooks, Ron. (2013). The Dream of the Thylacine. London : Frances Lincoln [distributor].

Xi, Shen Shi. (2010). Black Swan Amethyst(Chinese Edition) : HEI TIAN E ZI SHUI JING. Linking Publishing Co. Ltd. 360 pp.

Many references were gleaned from here and here. A few from here also

Many other newspaper article citations can be viewed here.

Trove, an online archive of old Australian newspapers, can be searched for mentions of the thylacine. Here are just three possible suggestions of searches: Tiger

The description by Brehm (1861) was brought to my attention by The Quest for Thylacoleo forum member arca. It is indeed of historical significance.

The pdf file of (Ackman, 1976) was kindly supplied by Gareth Linnard.

The pdf file of (Sharland, 1938) was indirectly sourced via Gareth Linnard from Thylacine Man.

Zoochat member snowleopard pointed out (Block, 2017)

(Image by Surroundx)
"Artist unknown, date unknown (pre-2010). From the author's private stamp collection."
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